Publications

Nazir SA, Ashby B, Leeks A & van Dijk B

Stellbrink L et al.

Seppälä O & Ashby B

King KC, Saad-Roy CM, Coombs D & Ashby B (2026) Microbial Primer: Understanding the exponential growth of epidemics. Microbiology. 172:001723

Ashby B & Anderson A (2026) Guarding versus self-guarding in innate immunity. Evolution.

Kim YS & Ashby B (2026) Coevolutionary cycling in allele frequencies and the evolution of virulence. Evolution. 80:282-290.

Smith CA, Renegado S & Ashby B (2025) The evolution of parasite virulence in the presence of resistance-conferring defensive symbionts. J. Evol. Biol. 39:1050-1058.

Ashby B & Best A (2025) Adaptive dynamics. Reference Module in Life Sciences

Smith CA & Ashby B (2025) Efficient coupling of within- and between-host infectious disease dynamics. J. Theor. Biol. 602-603:112061.

Wood J & Ashby B (2024) Diagnostic testing and the evolution of detection avoidance by pathogens. Evol. Med. Pub. Health. 12:248–259.

Buckingham LJ & Ashby B (2024) Coevolution of age-structured tolerance and virulence. Bull. Math. Biol. 86:62

Buckingham LJ & Ashby B (2024) Separation of Evolutionary Timescales in Coevolving Species. J. Theor. Biol. 579:111688.

Wood J & Ashby B (2023) Hyperparasitism and the evolution of parasite virulence. Evolution. 77:2631–2641.

Smith CA & Ashby B (2023) Tolerance-conferring defensive symbionts and the evolution of parasite virulence. Evol. Lett. 4:262-272.

Ashby B, Smith CA & Thompson RN (2023) Non-pharmaceutical interventions and the emergence of pathogen variants. Evol. Med. Pub. Health. 11:80-89.

Smith CA & Ashby B (2023) Antigenic evolution of SARS-CoV-2 in immunocompromised hosts. Evol Med Pub Health. 11:90-100.

Best A & Ashby B (2023) How do fluctuating ecological dynamics impact the evolution of hosts and parasites? Phil Trans R Soc B. 378: 20220006.

Buckingham LJ & Ashby B (2023) The evolution of the age of onset of resistance to infectious disease. Bull. Math. Biol. 85:42

Buckingham LJ, Bruns EL & Ashby B (2023) The evolution of age-specific resistance to infectious disease. Proc. R. Soc. B. 290:20222000

Höckerstedt L*, Numminen E*, Ashby B*, Boots M, Norberg A & Laine AL (2022) Spatially structured eco-evolutionary dynamics in a host-pathogen interaction render isolated populations vulnerable to disease. Nat Comm. 13:6018

Smith CA, Yates CA & Ashby B (2022) Critical weaknesses in shielding strategies for COVID-19. PLOS Glob Public Health 2:e0000298.

Ashby B & Farine D (2022) Social information use shapes the coevolution of sociality and virulence. Evolution. 76:1153-1169.

Kretzschmar ME, Ashby B, Fearon E, Overton CE, Panovska-Griffiths J, Pellis L, Quaife M, Rozhnova G, Scarabel F, Stage HB, Swallow B, Thompson RN, Tildesley MJ & Villela D (2022) Challenges for modelling interventions for future pandemics. Epidemics. 38:100546.

Buckingham LJ & Ashby B (2022). Coevolutionary theory of hosts and parasites. J. Evol. Biol. 35:205-224.

Pirrie A & Ashby B (2021). Does differential mortality after parental investment affect sex ratio evolution? No. Evolution. 75:3175-3180.

Halimubieke N*, Pirrie A*, Székely T & Ashby B. How do sex biases in demography and disease characteristics affect the spread of sexually transmitted infections? J. Theor. Biol. 527:110832.

Best A & Ashby B (2021) Evolutionarily stable strategies are well-studied in periodically fluctuating populations. Proc. Natl. Acad. Sci. USA. 118:e2102001118.

Ashby B & Best A (2021) Herd immunity. Curr. Biol. 31:R174-R177.

Hall AR, Ashby B, Bascompte J & King KC (2020) Measuring Coevolutionary Dynamics in Species-Rich Communities. Trends. Ecol. Evol. 35:539-550.

Ashby B (2020) When does parasitism maintain sex in the absence of Red Queen Dynamics? J. Evol. Biol. 33:1795-1805.

Thompson RN, Hollingsworth TD, Isham V, Arribas-Bel D, Ashby B, Britton T, Challenor P, Chappell LHK, Clapham H, Cunniffe NJ, Dawid AP, Donnelly CA, Eggo RM, Funk S, Gilbert N, Glendinning P, Gog JR, Hart WS, Heesterbeek H, House T, Keeling M, Kiss IZ, Kretzschmar ME, Lloyd AL, McBryde ES, McCaw JM, McKinley TJ, Miller JC, Morris M, O’Neill PD, Parag KV, Pearson CAB, Pellis L, Pulliam JRC, Ross JV, Scalia Tomba G, Silverman BW, Struchiner CJ, Tildesley MJ, Trapman P, Webb CR, Mollison D & Restif O (2020) Key Questions for Modelling COVID-19 Exit Strategies. Proc. R. Soc. B. 287:20201405.

Ashby B*, Shaw A* & Kokko H (2020) An inordinate fondness for species with intermediate dispersal abilities. Oikos. 129:311-319.

Featured as the Editor’s Choice article for this issue of Oikos.

Ashby B (2020) Antagonistic coevolution between hosts and sexually transmitted infections. Evolution. 74: 43-56.

Dimitriu T, Ashby B & Westra ER (2019) Transposition: A CRISPR way to get around. Curr. Biol. 29:R886-889.

Ashby B, Iritani R, Best A, White AR & Boots M (2019) Understanding the role of eco-evolutionary feedbacks in host-parasite coevolution. J. Theor. Biol. 464:115-125.

Ashby B & Bruns E (2018) The evolution of juvenile susceptibility to infectious disease. Proc. R. Soc. B. 285:20180844.

Rafaluk-Mohr C, Ashby B, Dahan D & King K (2018) Mutual fitness benefits arise during coevolution in a nematode-defensive microbe model. Evol. Lett. 2:246-256.

Bentkowski P, van Oosterhout C, Ashby B & Mock T (2017) The effect of extrinsic mortality on genome size evolution in prokaryotes. ISME. 11:1011-1018.

Best A, Ashby B, White A, Bowers R, Buckling A, Koskella B & Boots M (2017) Host–parasite fluctuating selection in the absence of specificity. Proc. R. Soc. B. 284:20171615.

Ashby B*, Watkins ER*, Lorenço J, Gupta S & Foster KR (2017) Competing species leave many potential niches unfilled. Nature Ecol. Evol. 1:1495-1501.

Ashby B & King KC (2017) Friendly foes: the evolution of host protection by a parasite. Evol. Lett. 1:211-221.

Ashby B & Boots M (2017) Multi-mode fluctuating selection in host–parasite coevolution. Ecology Letters. 20:357-365.

van Houte S, Ekroth A, Broniewski JM, Chabas H, Ashby B, Bondy-Denomy J, Gandon S, Boots M, Paterson S, Buckling A & Westra ER (2016) The diversity-generating benefits of a prokaryotic adaptive immune system. Nature. 532:385-388.

Ashby B & Boots M (2015) Coevolution of parasite virulence and host mating strategies. Proc. Natl. Acad. Sci. USA. 112:13290-13295.

Ashby B & King KC (2015) Diversity and the maintenance of sex by parasites. J Evol. Biol. 28:511-520.

Gómez P, Ashby B & Buckling A (2015) Population mixing promotes arms race host-parasite coevolution. Proc. R. Soc. B. 282:20142297.

Ashby B, Gupta S & Buckling A (2014) Effects of epistasis on infectivity range during host-parasite coevolution. Evolution. 68:2972-2982.

Ashby B & Gupta S (2014) Parasitic castration promotes coevolutionary cycling but also imposes a cost on sex. Evolution. 68:2234-2244.

Ashby B, Gupta S & Buckling A (2014) Spatial structure mitigates fitness costs in host-parasite coevolution. Am. Nat. 183: E64-E74.

Ashby B & Gupta S (2013) Sexually transmitted infections in polygamous mating systems. Philos. Trans. R. Soc. B. 368:20120048.

Penman BS, Ashby B, Buckee CO & Gupta S (2013) Pathogen selection drives nonoverlapping associations between HLA loci. Proc. Natl. Acad. Sci. USA. 110:19645-19650.

Ashby B (2009) Exploring children’s attitudes towards mathematics. Proc. Brit. Soc. Res. Learn. Math. 29:7-12.

Pirrie A., Chapman H. & Ashby B (2022) ‘Parasite-mediated sexual selection: to mate or not to mate?’ in Ezenwa V.O., Altizer, S. & Hall R. (ed.) Animal Behavior and Parasitism. Oxford Univ. Press, Oxford, UK. pp. 151-168

Ashby B, Jones J, Knell RJ & Hurst GDD (2019) ‘Sexually transmitted infections in natural populations: what have we learnt from beetles and beyond?’ in Wilson K, Fenton A, & Tompkins D. (ed.) Wildlife Disease Ecology: Linking Theory to data and application. Cambridge Univ. Press, Cambridge, UK. pp. 187-222.

Ashby B (2014) The effects of contact patterns and genetic specificity on host and parasite evolution.